Very very cool game! I really dig the pixel aesthetic. The gameplay reminds me of Echo the Dolphin, I loved that game as a kid. I agree with the other comment, some sort of pulse that guides you in the right direction would be much appreciated if you get stuck. The different modes is a really nice touch, I think its great to include things like that in a jam game.
My suggestion for how to develop it is go more into the metroidvania route, like Animal World where there's lots of cool items that unlock new paths and things to do. For a jam game tho, really excellent work one of my favourites i've played so far